|Types of Gladiolus|
Gladiolus are found in a variety of types that include both the species and hybrid glads. The different types of species represent the geographic and ecological range of the many species in this genus. The different combinations of species used to create the different hybrids has led to the establishment of several different types of hybrids as well.
The Cape region of South Africa is home to over 100 different species of Gladiolus. Most of the Cape species grow and bloom in the winter, which in this region of South Africa is mild, and similar to that of Southern California. Many of the species are reasonably hardy and will tolerate some frost. Unlike the more familiar grandiflora hybrids, these species are small plants, typically they produce one or just a few slender leaves before flowering. The flowers are small and bourne on slender stalks. Many of the species, such as Gladiolus tristis, pictured on the right, are strongly fragrant. Hybridizers in the late 19th and early 20th centuries produced many hybrids with these species. Virtually all gladiolus species are diploids with 30 chromosomes and are interfertile. Most of these hybrids are long gone, however more recent hybrids have been produced including the so-called Homoglads; a cross between Homoglossum watsonius (recently reclassified as Gladiolus watsonius) and Gladiolus tristis. Based on descriptions of some of these older hybrids, there is great potential for producing useful garden plants from these species.
I have also put together a page with my cultural techniques and observations. Winter flowering species cultural notes. Obtaining bulbs is something of a challenge in the US. There are a few growers of specialty bulbs that carry some species. Specialty seed companies and the International Bulb Society are sources for seeds. All of my current stock has come from Jim Duggan's Flower Nursery, 1452 Santa Fe Dr., Encinitas, CA 92024. (760) 943-1658, www.thebulbman.com. His stock is high quality and the bulbs have consistently bloomed the first year. His web site has an extensive collection of pictures of many different South African bulbs.
|Winter flowering species that I have grown and some notes:|
|Gladiolus caeruleus-A graceful, fairly vigourous plant growing to about 18" high when in bloom. Blooms for me in late January to early February (1/23/99, 2/3/00, & 1/25/01). Spikes have 6-8 buds and have two bell shaped light blue flowers in bloom at a time. The lower petals are cream colored and marked with many dark blue-violet spots, somewhat like an Alstromeria. The flower spike bends so that the flowers face down, probably as an adaptation to keep the pollen dry during wet periods. Each 1" diameter flower can last 4-5 days and has no scent. The plants are self-incompatible, but readily cross with other species (and presumably other plants of the same species). The plants will tolerate temperatures down to at least 27 F, but the flowers and buds can be damaged by below freezing temps. It likes plenty of moisture during the growing season. This species is one of the easiest to grow and flower, at least in my experience. It likes to be wet, but can tolerate drier conditions without triggering premature dormancy, and is not prone to rot if watered before it breaks dormancy in the fall. It also flowers from small bulbs and setting seed does not strain the plant too much, which happens with some other species.|
|Gladiolus carneus- A large flowered, late blooming species. The single spike had three buds with large light lavender pink florets. Each lightly ruffled floret had three prominent red-pink darts on the lip petals. The flowers had little or no fragrance. Set seed when pollinated with G. tristis.|
|Gladiolus huttonii- This species produces a single slender leaf from which a graceful spike emerges. The spike had 4 buds and had all four flowers open at once. The species has no detectable fragrance. Set seed abundantly when pollinated by G. tristis. Bloomed 3/8/00.|
|Gladiolus miniatus- A vigorous late spring blooming species (4/27/01). G. miniatus produces 10-12 bud spikes with medium sized light pink flowers. The flowers last about four days and the color deepens over time. Each flower has throat darts similar, but not as conspicous as G. carneus. In some ways it resembles a larger version of G. carneus, but lacks the fragrance characteristic of G. carneus.|
|Gladiolus priorii- A very early blooming, almost leafless species. If it is going to bloom, it produces a sheath and flower stalk right from the start of growth. If not, it will produce long flat leaves. Has bloomed twice for me on 12/22/98 and 12/24/00, with no bloom in the 1999-2000 season. In 1998, the spike had two buds, both long lasting (9 days) bright red florets were in bloom at once. In 2000, the spike had 4 buds and all four flowers opened in two days, each flower again lasted 9 days. The flowers have no scent, are somewhat spidery in form and have no fragrance. The plant did produce a few seeds from self-pollination.|
|Gladiolus teretifolius- A graceful and very attractive species. The zygomorphic bright red flowers look like butterflies. No fragrance, as is typical with the red flowered species. Flowered 3/1/00 and each flower lasted 6-7 days.|
|Gladiolus trichonemifolius- A small delicate species with yellow flowers and a freesia-like fragrance. It bloomed on 2/6/99 and produced a spike with a single bud. The light yellow flower was unlike the other Gladiolus species I've grown, in that the flower opened and closed according to daylight. Like some other Iridaceae from the Cape (Sparaxis grandiflora, Geissorhiza splendidissima and Moraea loubseri and neopavonia) and most cactus, the flower only opened in bright sunlight and closed if it became cloudy. Although this species likes water during growth, it easily rots if watered before it begins growth.|
|Gladiolus tristis- A hardy easy growing species. Produces a single leaf from which the flower spike emerges. Typically 4-6 buds per spike. The flowers are intensely fragrant, especially at night. The fragrance is similar in scent and intensity to that of a "Stargazer" lily. G. tristis is pictured at the top of the page. It is not self fertile, but sets seed readily when crossed with other species. G. tristis likes to be wet when in active growth, and will go dormant if drought stressed even for a very short time. If this happens, it will abort the flower spike and die back; continued watering will cause rot when the plants are going dormant. Bloomed 3/12/00|
|Gladiolus watsonius- A small plant producing a single round leaf. The flower spike emerges from the base of the leaf. This species is a rather vigourous, if dainty plant and in the second year has 6-8 bulblets sprouting around the base of the main bulb. It blooms in early to mid February (2/15/00 and 2/5/01) and has 2-3 buds. Each flower lasted 6-7 days. The flowers are burnt orange in color with the center of the flower, especially the "hood" petal being a pale yellow with some veining of the orange color. The flowers have no scent, are spidery in form and span 3" in diameter. They are strongly zygomorphic.|
In addition to the concentration of winter blooming species in the Cape region, many more species are found in the inland areas of South Africa and tropical Africa. Several species are even native to Europe. One of these summer-blooming African species, Gladiolus dalenii, is chiefly responsible for the production of the summer blooming hybrids. G. dalenii occurs over a large region in Southern Africa and has many distinct geographic forms, which in the past were given many different species names, including G. primulinus, a form that was very important in the production of miniature Gladiolus hybrids. Several other summer blooming species also figure in the development of the modern hybrids, including G. cruentus, G. oppositiflorus, G. papilio and G. saundersii. Many other species were grown at the time the early hybrids that lead to modern Gladiolus hybrids, but it is not clear what other species contributed to the modern germplasm. The genes of G. papilio are clearly present in modern glads as evidence by the blotched varieties. Either or both G. cruentes and G. saundersii contributed the red color found in many modern cultivars. For the most part these species are not readily available. However, several of the European species can be found in bulb catalogs. One species, Gladiolus muriale (formerly Acidanthera bicolor var Muriale) is commonly found in catalogs and garden centers, often under the name "peacock lily". G. muriale, which is pictured at left, is strongly scented and blooms very late in the summer, often just before frost in Northern areas. It has been used to develop several hybrids with modern Gladiolus, the best known is 'Lucky Star'. Unfortunately the fragrance in the hybrids is not as strong as in the species, and is readily lost in further crosses to the grandiflora type hybrids, and in fact 'Lucky Star' was the only fragrant seedling produced from a large number of crosses made by Mrs. Joan Wright in the 1950's between G. muriale and modern hybrids
The nanus group of hybrids is commonly found in catalogs and garden centers labeled as "Butterfly Glads" or "Winter-hardy Glads". Both of these names fit this group well. For a group of hybrids derived from African species, they are surprisingly hardy (to USDA zone 4 when overwintered in the ground and can also be treated like the Grandiflora hybrids and dug each fall and replanted in the spring). Many of the hybrids found today were bred at the turn of the century, which is a testament to the disease resistance of this group. Although the exact parentage of these hybrids is mostly lost, we know that they are the result of crossing the summer and winter blooming species together. In fact, virtually all of them are actually first generation hybrids. We know this because they are sterile, with three sets of chromosomes (triploid) instead of the normal two sets (diploid) or the four sets found in the modern Grandiflora hybrids. Because odd sets of chromosomes cannot pair during the first division of meiosis, normal gamets cannot be formed. However, in gladiolus hybrids, it has been found that triploids do produce some gamets and that these typically have either 15, 30 or 45 chromosomes. These viable gametes typically form in the ovaries and thus the plants are pollen sterile. If the triploid plants are pollinated by pollen from a diploid (15n) or tetraploid (30n), some of the resulting seeds will be fertile tetraploids (2n=60). The fact that most nanus glads are triploids indicates that they derive from crosses with G. dalenii, a Summer blooming species with four sets of chromosomes (tetraploid), and some of the many Winter blooming, diploid species.
The nanus glads display a variety of colors and patterns, but most have conspicuous darts of color on the upper petals. This is in contrast to the blotch pattern in the modern Summer blooming glad hybrids, which is on the lower petals. However, this pattern of darts is very similar to that seen in G. carneus. The nanus glads come in white, pink, salmon and some varieties are nearly red. They have graceful spikes with fewer than 12 buds. Because of their grace and hardiness they are useful garden subjects, and are grown as cutfloers in Europe. Recently, breeders in Israel have produced a new series of hybrids by crossing modern Summer flowering hybrids with the Winter blooming species G. tristis. These have been produced for the cutflower trade and should be similar in growth habit and bloom season to the nanus cultivars.
The modern Summer blooming hybrids are very different from the species glads. The modern hybrids or "Grandiflora" hybrids are much larger, both in terms of flower size and thesize of the flower spike. Some modern hybrids can have up to 40 flower buds and can hold ten or more 5 1/2" wide flowers open at once. Hydrids such as Alpine Sunset (shown on the right) have been bred for exhibition at the many gladiolus shows around the country. Many other hybrids have been bred for commericial trade. While typically not capable of producing the individual spikes that the exhibition varieties can, the commercial varieties can be cut in tight bud and open days later in water. These spikes can bloom for more than a week with proper treatment. The ease of handling and the brilliant colors make gladioli one of the premier cutflowers.
In addition to the larger size, modern hybrid glads have flowers in colors and forms not found in the species. Most modern glads have at least some ruffling of the petals and some varieties are so heavily ruffled and textured that they almost appear to be carved from wax. The hybrid glads can be found in virtually all colors from pure white to nearly jet black. Pink and salmon colored varieties are perhaps the most common, but rich reds, golden yellows, purples and browns are also found among the modern cultivars. Even green, blue and violet glads exist. Although these three colors are not "true" colors in glads, modern hybridizers have come a long way towards producing glads that really look green and blue. Part of the reason for the many colors and forms that modern glads take is in the number and kinds of species used to create them.
Although the vast majority of Gladiolus species are diploids with 30 chromosomes, the Grandiflora hybrids are all tetraploids with 60 chromosomes. The tetraploid nature of modern garden glads traces to the prominence of G. dalenii in the parentage of the modern hybrids. Unlike most other species, G. dalenii is a tetraploid with 60 chromosomes. G. dalenii occurs in a large number of different forms, and has a wide natural range. It is found in the interior of South Africa, and ranges north to central Africa. Because of its wide range and variability its many forms were originally described as separate species (G. psittacinus, G. dracocephalus, and G. natalensis from South Africa; G. primulinus and G.quartinianus from Tropical East Africa). Crosses between the diploid summer and winter blooming species and forms of G. dalenii resulted in many different hybrids. Although most of these hybrids were triploids and thus mostly sterile, in gladioli, triploids produce unreduced gametes at a high frequency. Because of this property, pollination of the triploids by diploid species results in a reasonable frequency of fertile tetraploid offspring. By this route, hybridizers at the turn of the century incorporated several other summer and winter blooming species, (including G. cardinalis, G. cruentes, G. oppositiflorus, G. papilio, and G. saundersii) leading to the creation of the modern Grandiflora hybrids.
Virtually all of these species were incorporated into the germplasm by the 1920's and virtually all of the improvement since that time has been produced by intercrossing hybrid varieties. In the 1940's, 50's and 60's gladioli reached the height of their diversity. Semi-double and fully double forms were bred along with lacinated forms and varieties with huge florets. Most of these varieties are now gone, however, the germplasm of today's gladioli certainly contains the capacity to reproduce these forms.
Even though today's varieties are not as diverse in form, hybridizers have greatly improved the degree of ruffling of the flowers and have continued to improve the number of buds produced per spike. Although many of the novelties of the past have disappeared, new variants such as the variety "Candy Cane" pictured here have been developed recently. Fragrant summer hybrids have been the subject of several modern hybridizer's efforts, but a strongly fragrant summer flowered hybrid with remains elusive.
The latest hybrids are unsurpassed in ruffling and color. In addition, the smaller flowered varieties have been highly developed, to the point that they rival the larger flowered varieties in form and style. Although most gladiolus varieties face forward, upward facing varieties, face-ups, have been developed. These varieties have long been known, but they have achieved something of a revival recently with ruffled varieties in new colors.
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